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Marcus Smith has been dropped to the bench by England head coach Steve Borthwick for Sunday's Six Nations clash vs Italy, with skipper Owen Farrell moving to fly-half as one of three changes.
To facilitate Farrell's shift from inside centre to No 10, Exeter Chiefs centre Henry Slade is brought into midfield after his recovery from a hip injury, as is Bath's Ollie Lawrence, with Joe Marchant dropped to the bench.
Among the forwards, Jack Willis is named to start in place of Ben Curry at openside flanker - the latter cut from the England squad earlier in the week, along with Ben Youngs and Anthony Watson.
Willis faces Italy at Twickenham two years on from suffering a serious knee injury in the same fixture, which kept him out of the game for 53 weeks.